Danny Jacobs
King Julien (voice) (13 eps)
Jeff Bennett
Horst / Hector / Willie / Sage (voice) (13 eps)
India de Beaufort
Clover / Crimson (voice) (13 eps)
Kevin Michael Richardson
Maurice (voice) (13 eps)
Andy Richter
Mort / Ted (voice) (13 eps)
Debra Wilson
Masikura (voice) (13 eps)
Betsy Sodaro
Xixi (voice) (13 eps)
Sarah Thyre
Becca / Dorothy (voice) (13 eps)
Diedrich Bader
Abner (voice) (9 eps)
David Krumholtz
Timo / Mom-Bot (voice) (7 eps)
JB Blanc
Mountain Lemur Captain (voice) (6 eps)
Dwight Schultz
Karl / Patrick (voice) (3 eps)
Henry Winkler
Uncle King Julien (voice) (3 eps)
David Koechner
Nurse Phantom (voice) (3 eps)
Grey DeLisle
Pam / Eloise (voice) (2 eps)
John Michael Higgins
Barty (voice) (2 eps)
Anjelica Huston
Julienne (voice) (2 eps)
Maurice LaMarche
Koto / Corpse (voice) (1 eps)
Kether Donohue
April (voice) (1 eps)
Lance Barber
Bruce (voice) (1 eps)
Max Greenfield
Kipper (voice) (1 eps)
Glen Powell
Trent (voice) (1 eps)
Frank Welker
Grandpa Mort (voice) (1 eps)
Bill Fagerbakke
King Joey (voice) (1 eps)
Crispin Freeman
Alex (voice) (1 eps)
Rob Paulsen
Brodney (voice) (1 eps)